At Fólkvangr, a front constructed inside an asteroid, the Gundam Lfrith is undergoing operating tests in the Vanadis Institute's laboratory. The Lfrith has not yet managed to meet the conditions imposed by the council, and test pilot Elnora Samaya is growing impatient. That same day, her daughter is celebrating her fourth birthday...

Substitute Soul Reaper Ichigo Kurosaki spends his days fighting against Hollows, dangerous evil spirits that threaten Karakura Town. Ichigo carries out his quest with his closest allies: Orihime Inoue, his childhood friend with a talent for healing; Yasutora Sado, his high school classmate with superhuman strength; and Uryuu Ishida, Ichigo's Quincy rival. Ichigo's vigilante routine is disrupted by the sudden appearance of Asguiaro Ebern, a dangerous Arrancar who heralds the return of Yhwach, an ancient Quincy king. Yhwach seeks to reignite the historic blood feud between Soul Reaper and Quincy, and he sets his sights on erasing both the human world and the Soul Society for good. Yhwach launches a two-pronged invasion into both the Soul Society and Hueco Mundo, the home of Hollows and Arrancar. In retaliation, Ichigo and his friends must fight alongside old allies and enemies alike to end Yhwach's campaign of carnage before the world itself comes to an end.

Princess Anisphia "Anis" Wynn Palletia has always dreamed of flying through the sky, even though the people of her kingdom consider it a silly ambition. Also at odds with her goal is the fact that Anis is incapable of using magic despite her noble status. Refusing to give up so easily, she renounces her right to the throne, and focuses on developing "magicology" by combining various resources with knowledge from her previous life on Earth. Due to Anis' nature, responsibility for the kingdom's future is passed onto her younger brother, Algard. Pushed into a political marriage with Euphyllia Magenta, a girl he hardly knows, Algard rebels by spending more time with a commoner girl than his fiancée. After Euphyllia tries to intervene, Algard publicly calls off their marriage and denounces his ex-fiancée as a bully. After crashing into the banquet where Algard makes the announcement, Anis rescues Euphyllia on the flying broom she is testing. She decides to make Euphyllia her assistant, which the other girl reluctantly agrees to. Although their partnership appears random at first, Anis has an ulterior motive for wanting Euphyllia's company.

During the summer holidays, energetic Hana Uzaki spent most of her time accompanying her lonesome upperclassman, Shinichi Sakurai. Now that school has resumed, Uzaki's teasing continues to ramp up, much to Sakurai's constant annoyance. Nevertheless, no amount of ridicule can damage the pair's relationship—which only seems to be getting better as their college days fly by!