Ever since moving to an in-game world after countless hours of VRMMO grinding, Yuna now calls Crimonia—the city where she started—home. While maintaining her reputation as a renowned adventurer, she continues to manage her businesses; but most importantly, Yuna spends time with the people she has grown to love. Nevertheless, this world has a lot more to offer the carefree adventurer. It is up to Yuna to discover these wonders, and perhaps help more people in need as she encounters their various plights.

Centuries ago, mankind abandoned planet Earth after the Vice, a race of mechanical aliens, drove them from their home into a life adrift in space. Now resigned to starships forged of pieces of Earth’s shattered moon, the final hope for humanity lies in the hands of Actresses, young women born with the ability to wield the only weapons that can harm the Vice: Alice Gears, mechanical suits that can finally turn the tide against the alien incursion.

It adapts the story from volume 12 (chapter 98 onwards) of the manga series "Demon Slayer: Kimetsu no Yaiba" . The story takes place in a new setting - Tanjiro arrives at the "Swordsmith Village." After the death of Upper Moon Six and Daki, a vacancy appears among the upper moons, and Muzan Kibutsuji summons the remaining upper moons to the Infinity Castle. Muzan is furious about the vacancy and believes that the upper moons should do everything in their power to find the "Blue Spider Lily" and eliminate the Demon Slayer Corps. As a result, Upper Moon Four, Hantengu, and Upper Moon Five, Gyokko, are dispatched to carry out the mission. Meanwhile, Tanjiro, who has been recovering from his injuries at the Butterfly Estate for two months, learns that Kozo Kanamori, the swordsmith, has refused to make him a new sword. Following the advice of the Hashira, Shinobu Kocho, Suma Nakahara, and Kiyo Terauchi, Tanjiro travels to the Swordsmith Village to conUpon arrival, he encounters the Love Hashira, Mitsuri Kanroji, and the Mist Hashira, Muichiro Tokito, as well as Genya Shinazugawa. Tanjiro and his companions are about to embark on a new battle. The opening theme song "Ketsui no Hikari" is performed by MAN WITH A MISSION and milet. The lyrics and composition are by Jean-Ken Johnny, with arrangement by MAN WITH A MISSION and cooperation by Chisato Ichinose. The ending theme song "Koikogare" is also performed by MAN WITH A MISSION and milet.

Movie adaptation of Tsurune: Kazemai Koukou Kyuudoubu.